Doctor of Philosophy in Biological/Doctor of Philosophy in Biomedical Sciences - Biochemistry, Quantitative Biology, Biophysics,Structural Biology

The Biochemistry, Quantitative Biology, Biophysics and Structural Biology (BQBS) Track provides students with experimental, theoretical, and computational research training across a broad range of scales, from molecules to cells to whole organisms. Our approaches encompass a full range of modern disciplines including biochemistry and molecular biology, biophysics, chemistry, cell biology, genomics and proteomics, computational biology, and structural biology. Recent revolutionary developments in modern biology have greatly advanced the diverse interests of our faculty. Technical advances in X-ray crystallography, cryo-electron microscopy (cryo-EM), live imaging, single molecule studies, next-generation sequencing, and mass spectrometry have led to a wealth of quantitative data for addressing long-standing biology questions. The integration of experimental data with computer modeling has stimulated productive collaborations among the disciplines of biology, chemistry, physics, and engineering. Our students will thus have a broad training in experimental and quantitative skills to perform in-depth mechanistic studies of diverse biological processes.

Because BBS students come from a variety of academic backgrounds and receive training at Yale in a wide array of disciplines, there are no common courses required of all BBS applicants. In general, successful applicants will have completed undergraduate coursework in biology, chemistry at least through organic chemistry, physics, and calculus. Each BBS Track may have more specific course requirements, which are described on the Track pages.Applicants must achieve a score of at least 600 on the paper version of the exam, 250 on the computer-based exam, and 100 on the internet-based exam.
The application will ask you to submit a Statement of Academic Purpose.
TOEFL - Applicants must achieve a score of at least 600 on the paper version of the exam, 250 on the computer-based exam, and 100 on the internet-based exam.